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Acknowledgements This work is dedicated to the loving memory of Professor Nir Friedman zla dear friend, a mentor, and a source of inspiration

Immunohistochemical staining in aged mice showed FOXO4 expression in Leydig cells, often with nuclear localization, whereas it was rarely expressed in young mice

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Tests in bacteria, cells, and mice all came back negative, suggesting the compound does not pose carcinogenic risk through DNA damage mechanisms